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Highly commended, behaviour: mammals
The great swim by Buddhilini de Soyza, Sri Lanka/Australia
Photograph: Buddhilini de Soyza/Wildlife Photographer of the Year/PA
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Highly commended, behaviour: birds
Up for grabs by Jack Zhi, US: In southern California, US, a juvenile white-tailed kite reaches to grab a live mouse from the clutches of its hovering father.
Photograph: Jack Zhi/Wildlife Photographer of the Year/PA
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Highly commended, urban wildlife
Lynx on the threshold by Sergio Marijuan, Spain: A young Iberian lynx pauses in the doorway of the abandoned hayloft where it was raised, on a farm in eastern Sierra Morena, Spain.
Photograph: Sergio Mariju/Wildlife Photographer of the Year/PA
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Highly commended, oceans – the bigger picture
Net loss by Audun Rikardsen, Norway: In the wake of a fishing boat, a slick of dead and dying herrings covers the surface of the sea off the coast of Norway.
Photograph: Audun Rikardsen/Wildlife Photographer of the Year/PA
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Highly commended, behaviour: amphibians and reptiles
The gripping end by Wei Fu, Thailand: Clutched in the coils of a golden tree snake, a red-spotted tokay gecko stays clamped onto its attacker’s head in a last attempt at defence.
Photograph: Wei Fu/Wildlife Photographer of the Year/PA
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Highly commended, plants and fungi
Mushroom magic by Juergen Freund, Germany/Australia
Photograph: Juergen Freund/Wildlife Photographer of the Year/PA
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Highly commended, natural artistry
Toxic design by Gheorghe Popa, Romania: A detail of a small river in the Geamana Valley, within Romania’s Apuseni Mountains
Photograph: Gheorghe Popa/Wildlife Photographer of the Year/PA
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Highly commended, animal portraits
Storm fox by Jonny Armstrong, US: A fox busy searching in the shallows for salmon carcasses.
Photograph: Jonny Armstrong/Wildlife Photographer of the Year/PA
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Highly commended, photojournalism
A caring hand by Douglas Gimesy, Australia: After a feed of special formula milk, an orphaned grey-headed flying-fox pup lies on a ‘mumma roll’, sucking on a dummy and cradled in the hand of a wildlife carer.
Photograph: Douglas Gimesy/Wildlife Photographer of the Year/PA
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Highly commended, animal portraits
Raw moment by Lara Jackson, UK
Photograph: Lara Jackson/Wildlife Photographer of the Year/PA
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Highly commended, wetlands – the bigger picture
The nurturing wetland by Rakesh Pulapa, India: Houses on the edge of Kakinada city reach the estuary, buffered from the sea by the remains of a mangrove swamp.
Photograph: Rakesh Pulapa/Wildlife Photographer of the Year/PA
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Highly commended, underwater
Deep feelers by Laurent Ballesta, France: A vibrant community of thousands of narwhal shrimps in deep water off the French Mediterranean coast.
Photograph: Laurent Ballesta/Wildlife Photographer of the Year/PA
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Highly commended, urban wildlife
Natural magnetism by Jaime Culebras, Spain: A tarantula hawk wasp clutching a tarantula in Quito, Ecuador.
Photograph: Jaime Culebras/Wildlife Photographer of the Year/PA
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Highly commended, 11-14 Years
Apollo landing by Emelin Dupieux, France: An Apollo butterfly settles on an oxeye daisy.
Photograph: Emelin Dupieux/Wildlife Photographer of the Year/PA
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Highly commended, 10 years and under
Lockdown chicks by Gagana Mendis Wickramasinghe, Sri Lanka: Three rose-ringed parakeet chicks pop their heads out of the nest hole as their father returns with food
Photograph: Gagana Mendis Wickramasinghe/Wildlife Photographer of the Year/PA
